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Jenner was born and grew up in Mt. Kisco, New York, the second of four children of William Jenner, a tree surgeon, and Estelle Jenner. William Jenner had competed in the U.S. Army Olympics in Nuremberg, Germany, in 1945, and won a silver medal in the 100-yard dash. In addition, Jenner's grandfather had run in several Boston Marathons. Jenner inherited their athletic ability and high energy level. Despite his athletic talent, Jenner soon grew to hate school because he had a reading disability that caused him to fail second grade (a big embarrassment). He would do anything to get out of reading in front of the class, and focused his energy on sports instead

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